Current Taxonomic Status of Late Cretaceous Ammonoid Species from the Western Interior Seaway Assigned to the Family Scaphitidae (Ancyloceratina: Scaphitaceae)

Jonathan R. Hendricks

Paleontological Research Institution, Ithaca, New York

Note: Names in green are accepted; names in red are synonyms. Return to main list here. This page was last updated October 13, 2017.

Class Cephalopoda

Order Ammonoidea Zittel, 1884

Suborder Ancyloceratina Wiedmann, 1966

Duration: Lower Tithonian to Upper Maastrichtian (Wright et al., 1996).

Superfamily Scaphitaceae Gill, 1871

Duration: Upper Albian to Maastrichtian (Wright et al., 1996).

Family Scaphitidae Gill, 1871

Duration: Uppler Albian to Maastrichtian (Wright et al., 1996).

PaleoDB taxon number: 95940.

Subfamily Scaphitinae Gill, 1871

Duration: Upper Albian to Upper Maastrichtian (Wright et al., 1996).

Genus Acanthoscaphites Nowak, 1911

Type species: Scaphites tridens Kner, 1850

Duration: Campanian (Wright et al., 1996).

PaleoDB taxon number: 14515.

Acanthoscaphites nodosus (Owen, 1852) = Hoploscaphites nodosus (Owen, 1852)

Reason for equivalency: moved to different genus (source: Landman et al., 2010).

Acanthoscaphites nodosus brevis (Meek, 1876) = Hoploscaphites brevis (Meek, 1876)

Reason for equivalency: synonym (source: Landman et al., 2010).

Acanthoscaphites nodosus quadrangularis (Meek and Hayden, 1860) = Hoploscaphites brevis (Meek, 1876)

Reason for equivalency: synonym (source: Landman et al., 2010).

Genus Discoscaphites Meek, 1871

Type species: Ammonites conradi Morton, 1834

Duration: Campanian to Maastrichtian (Wright et al., 1996).

PaleoDB taxon number: 14786.

Discoscaphites conradi (Morton, 1834)

Original presentation: Ammonites conradi

Original publication. Morton. 1834. [p. 39, pl. 16, fig. 3].

Holotype: ANSP 51551.

Type locality: awaiting determination.

Type horizon: Prairie Bluff Chalk.

Other specimens: YPM 27182, YPM 27104, YPM 27132, YPM 27185, YPM 27183, YPM 27188, YPM 27187, YPM 27186, YPM 27195, YPM 27129, YPM 27198, YPM 27128, YPM 27129, YPM 27106, YPM 27127, YPM 27133, YPM 27197, YPM 27196, YPM 27103, YPM 27120, YPM 27115, YPM 27112, YPM 27102, YPM 27111, YPM 34100, YPM 23048, AMNH 44232, AMNH 44677, USNM 468859; also see Landman and Waage (1993).

WIS occurrences:

Maastrichtian: Fox Hills Formation (Trail City Member) (ND)

Maastrichtian: Fox Hills Formation (Trail City Member) (SD)

Maastrichtian: Fox Hills Formation (Timber Lake Member)

Maastrichtian: Pierre Shale (Elk Butte Member)

Other occurrences: Prairie Bluff Chalk (AL), Kemp Clay (TX), Corsicana Marl (TX), Severn Fm. (MD) (Landman and Waage, 1993).

PaleoDB taxon number: 129476.

Synonyms: Discoscaphites roanensis Stephenson, 1941 (source: Landman and Waage, 1993).

Reference: Landman and Waage (1993).

Discoscaphites gulosus (Morton, 1834)

Original presentation: Ammonites conradi var. gulosus

Original publication: Morton. 1834. [p. 39, pl. 16, fig. 2].

Holotype: ANSP 51552.

Type locality: Alabama.

Type horizon: Prairie Bluff Chalk.

Other specimens: YPM 2086, YPM 27190, YPM 27190, USNM 468860, YPM 27110, YPM 27191,YPM 27200, YPM 27199, YPM 27193, YPM 27126, YPM 27114, YPM 27189, YPM 27202, YPM 27204, YPM 27194, YPM 27205YPM 27192, YPM 23736, YPM 27201, YPM 27203, YPM 27097, YPM 27121, YPM 27117, YPM 27124, YPM 27095, YPM 27110, YPM 34106, YPM 34107, YPM 23050, YPM 23051; also see Landman and Waage (1993).

WIS occurrences:

Maastrichtian: Fox Hills Formation (Trail City Member) (SD) (Landman and Waage, 1993).

Maastrichtian: Fox Hills Formation (Trail City Member) (ND) (Landman and Waage, 1993).

Maastrichtian: Fox Hills Formation (Timber Lake Member) (SD) (Landman and Waage, 1993).

Maastrichtian: Fox Hills Formation (Timber Lake Member) (ND) (Landman and Waage, 1993).

Other occurrences: Prairie Bluff Chalk (Alabama), Prairie Bluff Chalk (Mississippi), Corsicana Marl (Texas), Severn Formation (Maryland) (Landman and Waage, 1993).

PaleoDB taxon number: 129477.

Synonyms: Ammonites conradi var. petchialis Morton, 1834 (source: Landman and Waage, 1993); Ammonites conradi var. navicularis Morton, 1834 (source: Landman and Waage, 1993).

Reference: Landman and Waage (1993).

Discoscaphites nebrascensis (Owen, 1852) = Hoploscaphites nebrascensis (Owen, 1852)

Reason for equivalency: moved to different genus (source: Landman and Waage, 1993).

Discoscaphites nicolleti (Morton, 1842) = Hoploscaphites nicolleti (Morton, 1842)

Reason for equivalency: moved to different genus (source: Landman and Waage, 1993).

Discoscaphites roanensis Stephenson, 1941 = Discoscaphites conradi (Morton, 1834)

Reason for equivalency: junior synonym (source: Landman and Waage, 1993).

Discoscaphites rossi Landman and Waage, 1993

Original presentation: Discoscaphites rossi

Original publication: Landman, N. H. and K. M. Waage. 1993. Scaphitid ammonites of the Upper Cretaceous (Maastrichtian) Fox Hills Formation in South Dakota and Wyoming. Bulletin of the American Museum of Natural History, Number 215, 257 pp. [p. 231-240; figs. 181-189].

Holotype: YPM 23052.

Type locality: Dewey County, South Dakota.

Type horizon: Fox Hills Formation (Timber Lake Member).

Other specimens: AMNH 44340, AMNH 44335, AMNH 44336, AMNH 44344, AMNH 44679, AMNH 44680, YPM 27084, YPM 23085, YPM 23095, YPM 23108, YPM 23096, YPM 23090, YPM 23094, YPM 23097, YPM 27081, YPM 23100, YPM 23083, YPM 23105, YPM 23093, YPM 23103.

WIS occurrences:

Maastrichtian: Fox Hills Formation (Timber Lake Member (SD) (Landman and Waage, 1993).

PaleoDB taxon number: 145275.

Reference: Landman and Waage (1993).

Origin of name: “named in honor of Mrs. Helen Ross” (Landman and Waage, 1993, p. 232).

Genus Hoploscaphites Nowak, 1911

Type species: Ammonites constrictus J. Sowerby, 1817

Duration: Campanian to Upper Maastrichtian (Wright et al., 1996).

PaleoDB taxon number: 15015.

Hoploscaphites birkelundi Landman and Waage, 1993

Original presentation: Hoploscaphites birkelundi

Holotype: YPM 27172.

Type locality: Niobrara County, Wyoming.

Type horizon: Fox Hills Formation.

Other specimens: YPM 27210, YPM 27170, YPM 27181, YPM 27180, YPM 27169, YPM 27177, YPM 27179, YPM 27211, YPM 27218, YPM 27209, YPM 27210, YPM 27207, YPM 27208, YPM 27172, YPM 34698, YPM 27220, AMNH 44225; also see Landman and Waage (1993).

WIS occurrences:

Maastrichtian: Fox Hills Formation (WY) (Landman and Waage, 1993).

Zonation: Hoploscaphites birkelundi Zone, upper Maastrichtian (Cobban et al., 2006).

PaleoDB taxon number: 340269.

Reference: Landman and Waage (1993).

Origin of name: “named in honor of the late Dr. Tove Birkelund” (Landman and waage, 1993, p. 119).

Hoploscaphites brevis (Meek, 1876)

Original presentation: Scaphites nodosus var. brevis

Original publication: Meek. 1876. [p. 426, pl. 25, figs. 1a-c.]

Holotype: USNM 367.

Type locality: probably “south fork of the Cheyenne River in South Dakota” (Landman et al., 2010, p. 159).

Type horizon: unknown.

Other specimens: see Landman et al. (2010).

WIS occurrences:

Campanian: Baculites reesidei Zone (WIS) (Landman et al., 2010)

Campanian: Baculites cuneatus Zone (WIS) (Landman et al., 2010)

Campanian: Baculites compressus Zone (WIS) (Landman et al., 2010)

Campanian: Bearpaw Shale (Alberta) (Landman et al., 2010)

Campanian: Bearpaw Shale (MT) (Landman et al., 2010)

Campanian: Bearpaw Shale (Saskatchewan) (Landman et al., 2010)

Campanian: Didymoceras cheyennense Zone (WIS) (Landman et al., 2010)

Campanian: Livingston Formation (MT) (Landman et al., 2010)

Campanian: Pierre Shale (CO) (Landman et al., 2010)

Campanian: Pierre Shale (MT) (Landman et al., 2010)

Campanian: Pierre Shale (ND) (Landman et al., 2010)

Campanian: Pierre Shale (SD) (Landman et al., 2010)

Campanian: Pierre Shale (WY) (Landman et al., 2010)

Campanian: Pierre Shale (Weskan Shale Member) (KS) (Landman et al., 2010)

Campanian: Pierre Shale (Lake Creek Shale) (KS) (Landman et al., 2010)

PaleoDB taxon number: not in database.

Synonyms: Scaphites nodosus? var. Meek, 1859 (source: Landman et al., 2010); Scaphites nodosus var. quadrangulus Meek and Hayden, 1860 (source: Landman et al., 2010); Scaphites nodosus var. exilis Meek and Hayden, 1860 (source: Landman et al., 2010); Acanthoscaphites nodosus brevis (Meek, 1876) (source: Landman et al., 2010); Acanthoscaphites nodosus quadrangularis (Meek and Hayden, 1860) (source: Landman et al., 2010); Jeletzkytes brevis (Meek, 1876); Jeletzkytes nodosus brevis (source: Landman et al., 2010); Hoploscaphites landesi Riccardi, 1983 (source: Landman et al., 2010); see Landman et al. (2010) for complete synonymy.

References: Landman et al. (2010).

Hoploscaphites comprimus (Owen, 1852)

Original presentation: Scaphites (Ammonites?) comprimus

Original publication: Owen. 1852. [p. 580, pl. 7, fig. 4].

Holotype: University of Chicago (UC) 6380 (at FMNH).

Type locality: “Fox Hills, Nebraska” (Owen, 1852).

Type horizon: probably “exposures of the Timber Lake Member” of the Fox Hills Formation (Landman and Waage, 1993, p. 100).

Other specimens: USNM 407, USNM 408, USNM 409, USNM 20243, YPM 27233, YPM 27230, YPM 23741, YPM 27229, YPM 27232, YPM 27228, YPM 27066, YPM 27231, YPM 27242, YPM 27234, YPM 27240, YPM 27239, YPM 27241, YPM 27146; see Landman and Waage (1993).

WIS occurrences:

Maastrichtian: Fox Hills Formation (Timber Lake Member) (SD) (Landman and Waage, 1993).

Maastrichtian: Fox Hills Formation (Trail City Member) (SD) (Landman and Waage, 1993).

PaleoDB taxon number: 145260.

References: Landman and Waage (1993).

Hoploscaphites dorfi (Landman and Waage, 1993)

Original presentation: Jeletzkytes dorfi

Holotype: YPM 23175.

Type locality: Niobrara County, Wyoming.

Type horizon: Fox Hills Formation.

Other specimens: YPM 23176, YPM 23174, YPM 23177, YPM 23212, YPM 27215, YPM 23185, YPM 27216, YPM 23179, YPM 23179, YPM 23180, YPM 27214, YPM 23212, AMNH 44337, AMNH 44224, AMNH 44339; also see Landman and Waage (1993).

WIS occurrences:

Maastrichtian: Fox Hills Formation (WY) (Landman and Waage, 1993).

Maastrichtian: Pierre Shale (SD) (Landman and Waage, 1993).

PaleoDB taxon number: 145271.

Reference: Landman and Waage (1993).

Origin of name: “named in honor of the late Erling Dorf” (Landman and Waage, 1993, p. 184).

Hoploscaphites landesi Riccardi, 1983 = Hoploscaphites brevis (Meek, 1876)

Reason for equivalency: junior synonym (microconchs of Hoploscaphites brevis) (source: Landman et al., 2010).

Hoploscaphites melloi Landman and Waage, 1993

Original presentation: Hoploscaphites melloi

Holotype (macroconch): YPM 23044.

Type locality: “inner bluffs of the Moreau River, northeast of the village of Green Grass and one mile west of the S.D. Rt. 63 bridge” (Landman and Waage, 1993, p. 113), Dewey County, South Dakota.

Type horizon: Pierre Shale (Mobridge Member).

Other specimens: YPM 23043, YPM 23071, YPM 23066, YPM 23067, YPM 23042, YPM 23068, YPM 23074, YPM 23077, YPM 23070, YPM 23072; also see Landman and Waage (1993).

WIS occurrences:

Maastrichtian: Pierre Shale (Mobridge Member) (Landman and Waage, 1993).

PaleoDB taxon number: 145262.

Reference: Landman and Waage (1993).

Origin of name: “named for James F. Mellow, micropaleontologist at the U. S. National Museum” (Landman and Waage, 1993, p. 113).

Hoploscaphites nebrascensis (Owen, 1852)

Original presentation: Ammonites nebrascensis

Original publication: Owen. 1952. [figs. 6g-i, 7a-c, g, h, 8a-d, g-i, 9g-j].

Lectotype: USNM 20242.

Type locality: “Fox Hills, Nebraska” (Kennedy et al., 1998, p. 766).

Type horizon: likely “Timber Lake Member of the Fox Hills Formation” (Landman and Waage, 1993, p. 161).

Other specimens: USNM 398, USNM 402, UC 6379, YPM 23146, YPM 23165, YPM 23144, YPM 23145, YPM 23172, YPM 23151, YPM 23147, YPM 23147, YPM 23687, YPM 23198, YPM 23195, YPM 23697, YPM 23735, YPM 23735, MAPS A2046a; also see Landman and Waage (1993).

WIS occurrences:

Maastrichtian: Fox Hills Formation (Timber Lake Member) (SD) (Landman and Waage, 1993).

Maastrichtian: Fox Hills Formation (Timber Lake Member) (WY) (Landman and Waage, 1993).

Maastrichtian: Fox Hills Formation equivalent (SD) (Landman and Waage, 1993).

Maastrichtian: Fox Hills Formation equivalent (ND) (Landman and Waage, 1993).

Maastrichtian: Fox Hills Formation (NE) (Kennedy et al., 1998)

Maastrichtian: Pierre Shale (Elk Butte Member) (Kennedy et al., 1998).

Other occurrences: “Severn Formation of Maryland” (Landman and Waage, 1993, p. 161).

Zonation: Hoploscaphites nebrascensis zone (= Jeletzkytes nebrascensis zone), upper Maastrichtian (Cobban et al., 2006).

PaleoDB taxon number: 129481.

Synonyms: Scaphites conradi Morton, 1856 (source: Landman and Waage, 1993); Ammonites cheyennensis Owen, 1852 (source: Landman and Waage, 1993), Ammonites moreauensis Owen, 1852 (source: Landman and Waage, 1993).

References: Landman and Waage (1993); Kennedy et al. (1998); Landman et al. (2015).

Origin of name: Nebraska Territory (see Cobban et al., 2006).

Hoploscaphites nicolletii (Morton, 1842)

Original presentation: Ammonites nicolletii

Original publication. Morton. 1842. [p. 209, pl. 10, fig. 3].

Neotype: YPM 27222 (plaster cast of Morton’s type is ANSP 51563).

Type locality: unknown (see Landman and Waage, 1993).

Other specimens: YPM 27221, YPM 27224, YPM 23704, YPM 27226, YPM 27247, YPM 23041, YPM 27244, YPM 27248, YPM 27225, YPM 27246, YPM 27223, YP 27245, YPM 27235, YPM 23707, YPM 23742, YPM 27243, YPM 27227, YPM 27238, YPM 27237, YPM 27144, YPM 27236; see Landman and Waage (1993).

WIS occurrences:

Maastrichtian: Fox Hills Formation (Trail City Member) (SD) (Landman and Waage, 1993).

Maastrichtian: Fox Hills Formation (Trail City Member) (ND) (Landman and Waage, 1993).

Maastrichtian: Pierre Shale (Elk Butte Member) (SD) (Landman and Waage, 1993).

Zonation: Hoploscaphites nicolletii Zone, upper Maastrichtian (Cobban et al., 2006).

PaleoDB taxon number: 145232.

Synonyms: ?Ammonites mandanensis Morton, 1842 (source: Landman and Waage, 1993); ?Ammonites abyssinus Morton, 1842 (source: Landman and Waage, 1993).

References: Landman and Waage (1993).

Origin of name: named for Joseph Nicolas Nicollet, “who explored and mapped the upper Mississippi River Valley and collected Cretaceous fossils in South Dakota in 1839” (Cobban et al., 2006, p. 33).

Hoploscaphites nodosus (Owen, 1852)

Original presentation: Scaphites (Ammonites?) nodosus (Owen, 1852)

Original publication. Owen. 1852. [p. 581, pl. 8, fig. 4].

Holotype: UC 6381.

Type locality: probably “near Sage Creek, Pennington County, South Dakota” (Landman et al., 2010, p. 99).

Type horizon: “probably from the Baculites cuneatus Zone of the Pierre Shale” (Landman et al., 2010, p. 99).

Other specimens: see listings in Landman et al. (2010).

WIS occurrences:

Campanian: Baculites compressus Zone (Western Interior Seaway) (Landman et al., 2010)

Campanian: Baculites cuneatus Zone (Western Interior Seaway) (Landman et al., 2010)

Campanian: Didymoceras cheyennense Zone (Western Interior Seaway) (Landman et al., 2010)

Campanian: Baculites reesidei Zone (Western Interior Seaway) (Landman et al., 2010)

Campanian: Bearpaw Shale (MT) (Landman et al., 2010)

Campanian: Livingston Formation (MT) (Landman et al., 2010)

Campanian: Pierre Shale (MT) (Landman et al., 2010)

Campanian: Pierre Shale (ND) (Landman et al., 2010)

Campanian: Pierre Shale (SD) (Landman et al., 2010)

Campanian: Pierre Shale (WY) (Landman et al., 2010)

Campanian: Pierre Shale (CO) (Landman et al., 2010)

Campanian: Pierre Shale (Weskan Shale Member) (KS) (Landman et al., 2010)

Campanian: Pierre Shale (Lake Creek Member) (KS) (Landman et al., 2010)

PaleoDB taxon number: not in database.

Synonyms: Scaphites nodosus Owen, 1852 (source: Landman et al., 2010); Acanthoscaphites nodosus (Owen, 1852) (source: Landman et al., 2010); Jeletzkytes nodosus (Owen, 1852) (source: Landman et al., 2010); see Landman et al. (2010) for complete synonymy.

References: Landman et al. (2010).

Hoploscaphites spedeni (Landman and Waage, 1993)

Original presentation: Jeletzkytes spedeni

Holotype: YPM 27160.

Type locality: Corson County, South Dakota.

Type horizon: Fox Hills Formation (Trail City Member).

Other specimens: YPM 23710, YPM 23124, YPM 24817, YPM 27156, YPM 23122, YPM 23206, YPM 23211, YPM 23775, YPM 23778, YPM 23205, YPM 23704, YPM 23199, YPM 23718, YPM 23204, YPM 23699, YPM 23200, YPM 23706, YPM 23714, YPM 31803, YPM 27156, YPM 27153, YPM 27151, YPM 27152, YPM 23710, YPM 27091, YPM 23062, YPM 23061, YPM 32593, YPM 32602, USNM 468854, USNM 468855, USNM 468856, USNM 468857, USNM 468858; also see Landman and Waage (1993).

WIS occurrences:

Maastrichtian: Fox Hills Formation (Trail City Member) (ND) (Landman and Waage, 1993).

Maastrichtian: Fox Hills Formation (Trail City Member) (SD) (Landman and Waage, 1993).

Maastrichtian: Pierre Shale (Elk Butte Member) (ND) (Landman and Waage, 1993).

Maastrichtian: Pierre Shale (Elk Butte Member) (SD) (Landman and Waage, 1993).

PaleoDB taxon number: 145264.

Synonyms: Jeletzkytes spedeni (Landman and Waage, 1993).

Reference: Landman and Waage (1993).

Origin of name: “named for Ian G. Speden” (Landman and Waage, 1993, p. 129).

Genus Jeletzkytes Riccardi, 1983 = Hoploscaphites Nowak, 1911

Reason for equivalency: synonym (source: Wright et al., 1996; Landman et al., 2015).

Jeletzkytes brevis (Meek, 1876) = Hoploscaphites brevis (Meek, 1876)

Reason for equivalency: Jeletzkytes is a junior synonym of Hoploscaphites.

Jeletzkytes dorfi Landman and Waage, 1993 = Hoploscaphites dorfi (Landman and Waage, 1993)

Reason for equivalency: Jeletzkytes is a junior synonym of Hoploscaphites.

Jeletzkytes nebrascensis (Owen, 1852) = Hoploscaphites nebrascensis (Owen, 1852)

Reason for equivalency: Jeletzkytes is a junior synonym of Hoploscaphites.

Jeletzkytes nodosus (Owen, 1852) = Hoploscaphites nodosus (Owen, 1852)

Reason for equivalency: Jeletzkytes is a junior synonym of Hoploscaphites.

Jeletzkytes nodosus brevis (Meek, 1876) = Hoploscaphites brevis (Meek, 1876)

Reason for equivalency: Jeletzkytes is a junior synonym of Hoploscaphites.

Jeletzkytes spedeni Landman and Waage, 1993 = Hoploscaphites spedeni (Landman and Waage, 1993)

Reason for equivalency: Jeletzkytes is a junior synonym of Hoploscaphites.

Genus Scaphites Parkinson, 1811

Type species: Scaphites equalis J. Sowerby, 1813

Duration: Upper Albian to Campanian (Wright et al., 1996).

Scaphites aquilaensis Reeside, 1927 = Scaphites hippocrepis I, II (DeKay, 1828)

Reason for equivalency: junior synonym (source: Cobban, 1969).

Scaphites aquilaensis var. costatus Reeside, 1927 = Scaphites hippocrepis I (DeKay, 1828)

Reason for equivalency: junior synonym (source: Cobban, 1969).

Scaphites aquilaensis var. nanus Reeside, 1927 = Scaphites hippocrepis I (DeKay, 1828)

Reason for equivalency: junior synonym (source: Cobban, 1969).

Scaphites aquisgranensis Schlüter, 1876 = Scaphites hippocrepis III (DeKay, 1828)

Reason for equivalency: junior synonym (source: Cobban, 1969).

Scaphites (Ammonites?) comprimus (Owen, 1852) = Hoploscaphites comprimus (Owen, 1852)

Reason for equivalency: moved to different genus (source: Landman and Waage, 1993).

Scaphites cuvieri Morton, 1876 = Scaphites hippocrepis III (DeKay, 1828)

Reason for equivalency: junior synonym (source: Cobban, 1969).

Scaphites levis Reeside, 1927 = Scaphites hippocrepis I (DeKay, 1828)

Reason for equivalency: moved to different genus (source: Landman and Waage, 1993).

Scaphites hippocrepis I (DeKay, 1828)

Original presentation: Ammonites hippocrepis

Original publication: DeKay. 1827. [p. 273, pl. 5, fig. 5].

Holotype: lost (see Cobban, 1969).

Type locality: Delaware.

Type horizon: Merchantville Formation.

Other specimens: USNM 160250-160291.

WIS occurrences:

Campanian: Cody Shale (WY) (Cobban, 1969)

Campanian: Telegraph Creek Formation(Elk Basin Sandstone Member) (WY) (Cobban, 1969)

Campanian: Eagle Sandstone (MT) (Cobban, 1969)

Campanian: Mancos Shale (NM) (Cobban, 1969)

Zonation: Scaphites hippocrepis I Zone, lower Campanian (Cobban et al., 2006)

PaleoDB taxon number: 324058.

Synonyms: Scaphites aquilaensis Reeside, 1927 [part] (source: Cobban, 1969); Scaphites aquilaensis var. costatus Reeside, 1927 (source: Cobban, 1969); Scaphites aquilaensis var. nanus Reeside, 1927 (Cobban, 1969); Scaphites levis Reeside, 1927 (source: Cobban, 1969).

References: Cobban (1969); Gill and Cobban (1973).

Origin of name: Greek “hippos” =horse + Greek “krepis” =shoe (Cobban, 1969).

Scaphites hippocrepis II (DeKay, 1928)

Original presentation: Ammonites hippocrepis

Original publication: DeKay. 1827. [p. 273, pl. 5, fig. 5].

Holotype: lost (see Cobban, 1969).

Type locality: Delaware.

Type horizon: Merchantville Formation.

Other specimens: USNM 160292-160301, USNM 160304, USNM 160305.

WIS occurrences:

Campanian: Eagle Sandstone (MT) (Cobban, 1969)

Campanian: Cody Shale (WY) (Cobban, 1969)

Zonation: Scaphites hippocrepis II Zone, lower Campanian (81.86 +/- 0.36 Ma) (Cobban et al., 2006)

PaleoDB taxon number: 324058.

Synonyms: Scaphites aquilaensis Reeside, 1927 (source: Cobban, 1969); Scaphites hippocrepis var. tenuis Reeside, 1927 (source: Cobban, 1969); Scaphites stantoni Reeside, 1927 (source: Cobban, 1969)

References: Cobban (1969); Gill and Cobban (1973).

Origin of name: Greek “hippos” =horse + Greek “krepis” =shoe (Cobban, 1969).

Scaphites hippocrepis III (DeKay, 1928)

Original presentation: Ammonites hippocrepis

Original publication: DeKay. 1827. [p. 273, pl. 5, fig. 5].

Holotype: lost (see Cobban, 1969).

Type locality: Delaware.

Type horizon: Merchantville Formation.

Other specimens: USNM 131465, USNM 160306-160334.

WIS occurrences:

Campanian: Cody Shale (WY) (Cobban, 1969)

Campanian: Pierre Shale (NM) (Cobban, 1969)

Campanian: Pierre Shale (Gammon Ferruginous Member) (WY) (Cobban, 1969)

Campanian: Pierre Shale (Gammon Ferruginous Member) (SD) (Cobban, 1969)

Campanian: Mancos Shale (UT) (Cobban, 1969)

Other occurrences: Atlantic and Gulf Coastal Plains, including TX, AL, GA, NJ, and DE (see Cobban, 1969, p. 24); Germany, France, Netherlands, and possibly Sweden (see Cobban, 1969, p. 24).

Zonation: Scaphites hippocrepis III Zone (Cobban et al., 2006).

PaleoDB taxon number: 324058.

Synonyms: Scaphites aquisgranensis Schlüter, 1876 (source: Cobban, 1969); Scaphites cuvieri Morton, 1876 (source: Cobban, 1969); Scaphites reniformis Morton, 1834 (source: Cobban, 1969); Scaphites similis Whitfield, 1892 (source: Cobban, 1969).

References: Cobban (1969); Gill and Cobban (1973).

Origin of name: Greek “hippos” =horse + Greek “krepis” =shoe (Cobban, 1969).

Scaphites hippocrepis var. tenuis Reeside, 1927 = Scaphites hippocrepis II (DeKay, 1928)

Reason for equivalency: junior synonym (source: Cobban, 1969).

Scaphites leei I Reeside, 1927

Original presentation: Scaphites leei

Original publication: Reeside. 1927.

Holotype: USNM 73354.

Type locality: Santa Fe County, New Mexico (Cobban, 1969).

Type horizon: Mancos Shale (Cobban, 1969).

Other specimens: USNM 160231-160233.

WIS occurrences:

Turonian to Santonian: Marias River Shale (MT) (Cobban, 1969)

Niobrara Shale (MT) (Cobban, 1969)

Mancos Shale (NM) (Cobban, 1969)

PaleoDB taxon number: 341399.

Reference: Cobban (1969).

Scaphites leei II Reeside, 1927

Original presentation: Scaphites leei

Original publication: Reeside. 1927.

Holotype: USNM 73354.

Type locality: Santa Fe County, New Mexico (Cobban, 1969).

Type horizon: Mancos Shale (Cobban, 1969).

Other specimens: USNM 160234-160236.

WIS occurrences:

Mancos Shale (NM) (Cobban, 1969)

Telegraph Creek Formation (MT) (Cobban, 1969)

Gammon Shale (MT) (Cobban, 1969)

PaleoDB taxon number: 341399.

Reference: Cobban (1969).

Scaphites leei III Reeside, 1927

Original presentation: Scaphites leei

Original publication: Reeside. 1927.

Holotype: USNM 73354.

Type locality: Santa Fe County, New Mexico (Cobban, 1969).

Type horizon: Mancos Shale (Cobban, 1969).

Other specimens: USNM 160237-160249.

WIS occurrences:

Campanian: Gammon Shale (MT) (Cobban, 1969)

Campanian: Telegraph Creek Formation (MT) (Cobban, 1969)

Campanian: Cody Shale (WY) (Cobban, 1969)

Campanian: Austin Chalk (TX) (Cobban, 1969)

Zonation: Scaphites leei III Zone, lower Campanian (Cobban et al., 2006).

PaleoDB taxon number: 341399.

Synonyms: Scaphites leei var. parvus Reeside, 1927 (source: Cobban, 1969).

Reference: Cobban (1969).

Scaphites leei var. parvus Reeside, 1927 = Scaphites leei III Reeside, 1927

Reason for equivalency: junior synonym (source: Cobban, 1969).

Scaphites mandanensis (Morton, 1842) = Hoploscaphites nicolletii (Morton, 1842)

Reason for equivalency: synonym (questionable microconch of H. nicolletii) (source: Landman and Waage, 1993).

Scaphites nicolletii (Morton, 1842) = Hoploscaphites nicolletii (Morton, 1842)

Reason for equivalency: moved to different genus (source: Landman and Waage, 1993).

Scaphites nodosus Owen, 1852 = Hoploscaphites nodusus (Owen, 1852)

Reason for equivalency: moved to different genus (source: Landman et al., 2010).

Scaphites nodosus? var. Meek, 1859 = Hoploscaphites brevis (Meek, 1876)

Reason for equivalency: synonym (source: Landman et al., 2010).

Scaphites nodosus var. exilis Meek and Hayden, 1860 = Hoploscaphites brevis (Meek, 1876)

Reason for equivalency: synonym (source: Landman et al., 2010).

Scaphites nodosus var. quadrangulus Meek and Hayden, 1860 = Hoploscaphites brevis (Meek, 1876)

Reason for equivalency: synonym (source: Landman et al., 2010).

Scaphites reniformis Morton, 1834 = Scaphites hippocrepis III (DeKay, 1928)

Reason for equivalency: junior synonym (source: Cobban, 1969).

Scaphites similis Whitfield, 1892 = Scaphites hippocrepis III (DeKay, 1928)

Reason for equivalency: junior synonym (source: Cobban, 1969).

Scaphites stantoni Reeside, 1927 = Scaphites hippocrepis II (DeKay, 1928)

Reason for equivalency: junior synonym (source: Cobban, 1969).

Genus Trachyscaphites Cobban and Scott, 1964

Type species: Trachyscaphites redbirdensis Cobban and Scott, 1964

Duration: Campanian (Wright et al., 1996).

Trachyscaphites densicostatus Cobban and Kennedy, 1992

Original presentation: Trachyscaphites densicostatus

Original publication: Cobban, W. A. and W. J. Kennedy. 1992. Campanian Trachyscaphites spiniger ammonite fauna in north-east Texas. Palaeontology 35(1): 63-93.

Holotype: USNM 441426.

Type locality: USGS Mesozoic locality D10154, Fannin County, Texas (Cobban and Kennedy, 1992).

Type horizon: Ozan Formation.

Other specimens: USNM 441427, USNM 441428.

WIS occurrences:

Campanian: Ozan Formation (TX) (Cobban and Kennedy, 1992).

PaleoDB taxon number: 343907.

Reference: Cobban and Kennedy, 1992.

Trachyscaphites spiniger (Schlüter, 1872)

Original presentation: Scaphites spiniger

Original publication: Schlüter. 1872. [p. 82, pl. 25, figs. 1-7].

Lectotype: “original [figure] of Schlüter 1872, pl. 25, figs 1-3” (Cobban and Kennedy, 1992, p. 86); designated by Błazkiewicz (1980).

Type locality: Darup, Westphalia, Germany.

Type horizon: “upper Campanian” (Cobban and Kennedy, 1992, p. 86).

Other specimens: GPIB 61a (paralectotype); GPIB 61b (paralectotype), USNM 441416-441425.

WIS occurrences:

Campanian: Ozan Formation (TX) (Cobban and Kennedy, 1992).

Campanian: Annona Chalk (TX) (Cobban and Kennedy, 1992).

Other occurrences: “Germany, The Netherlands, Sweden, Poland, the USSR” (Cobban and Kennedy, 1992, p. 88).

PaleoDB taxon number: 130781.

References: Cobban and Kennedy (1992).